I’m wondering if extremely low water (like dry, or at least non continuous flow) conditions on the tribs forced the Brookies DOWN into Penns during the extremely low flow Summers we’ve recently had. I suspect Swift and Panther may have seen conditions like that. Cherry, Weikert, and Poe are a good bit bigger, but still possible. I found non-continuous flow on Hammersley Fork, mid-watershed, the last few years. Just a thought. I don’t know, just another potential angle to consider. They got down there, found suitable, or at least survivable conditions, and stayed?
Definitely some streams in the Loyalsock drainage that were completely dry for significant lengths the last few years.